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It depends on the pattern. Ingrid's explanation is the correct one for the twist sts but if your pattern has defined the sts differently, then that should be what is required.
I don't see any complaints in glancing at the Ravelry page and there aren't errata online.
The pattern for the larger branches looks like it takes 3 sts while the smaller twigs look like a 2 stitch twist. Which ones are you doing?